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J. Beddington (1974)
Age Structure, Sex Ratio and Population Density in the Harvesting of Natural Animal PopulationsJournal of Applied Ecology, 11
R. Law (2000)
Fishing, selection, and phenotypic evolutionIces Journal of Marine Science, 57
(1966)
Elgen i Norge sett med jegerøyne. En analyse av jaktmaterialet fra overv˚akningsprogrammet for elg og det samlede Sett-elg materialet for perioden
E. Pollak (2000)
The effective population size of some age-structured populations.Mathematical biosciences, 168 1
E. Solberg, C. Rolandsen, Morten Heim, Vidar Grøtan, Mathieu Garel, Bernt-Erik Sæther, E. Nilsen, Gunnar Austrheim, Ivar Herfindal (2006)
Elgen i Norge sett med jegerøyne. En analyse av jaktmaterialet fra overvå-kingsprogrammet for elg og det samlede sett elg-materialet for perioden 1966-2004
J. Beddington, D. Taylor (1973)
356 NOTE: Optimum Age Specific Harvesting of a PopulationBiometrics, 29
Mandy Haldane (1927)
A Mathematical Theory of Natural and Artificial Selection, Part V: Selection and MutationMathematical Proceedings of the Cambridge Philosophical Society, 23
Ø. Holand, K. Røed, A. Mysterud, J. Kumpula, M. Nieminen, Martin Smith (2003)
The effect of sex ratio and male age structure on reindeer calvingJournal of Wildlife Management, 67
B. Sæther, E. Solberg, Morten Heim (2003)
Effects of Altering Sex Ratio Structure on the Demography of an Isolated Moose PopulationJournal of Wildlife Management, 67
L. Nunney (1991)
The influence of age structure and fecundity on effective population sizeProceedings of the Royal Society of London. Series B: Biological Sciences, 246
(1998)
Social organization and effective population sizes in carnivores
Cedric Smith, S. Wright (1968)
Evolution and the Genetics of Populations: Volume 2, The Theory of Gene Frequencies
Tonje Stubsjøen, B. Sæther, E. Solberg, Morten Heim, C. Rolandsen (2000)
Moose (Alces alces) survival in three populations in northern NorwayCanadian Journal of Zoology, 78
M. Garel, Jean, Marc Cugnasse, D. Maillard, Michel Gaillard, J. A., M. Hewison, D. Dubray (2007)
Selective harvesting and habitat loss produce long-term life history changes in a mouflon population.Ecological applications : a publication of the Ecological Society of America, 17 6
G. (2003)
A NOTE ON EFFECTIVE POPULATION SIZE WITH OVERLAPPING GENERATIONS
E. Solberg, A. Loison, T. Ringsby, B. Sæther, Morten Heim (2002)
Biased adult sex ratio can affect fecundity in primiparous moose Alces alcesWildlife Biology, 8
(1987)
Pattern and processes in the population dynamics of Scandinavian moose ( Alces alces ) : some suggestions
Beddington Beddington, Taylor Taylor (1973)
Optimum age specific harvesting of a populationBiometrics, 29
Ginsberg Ginsberg, Milner‐Gulland Milner‐Gulland (1994)
Sex‐biased harvesting and population dynamics in ungulatesimplications for conservation and sustainable use, 8
W. Hill (1979)
A note on effective population size with overlapping generations.Genetics, 92 1
S. Engen, R. Lande, Bernt-Erik aether, H. Weimerskirch (2005)
Extinction in relation to demographic and environmental stochasticity in age-structured models.Mathematical biosciences, 195 2
Lee Lee, Engen Engen, Sæther Sæther (2008)
Understanding mating systemsa mathematical model of the pair formation process, 73
Fenberg Fenberg, Roy Roy (2008)
Ecological and evolutionary consequences of size‐selective harvestinghow much do we know?, 17
A. Caballero, W. Hill (1992)
Effective size of nonrandom mating populations.Genetics, 130 4
Røed Røed, Holand Holand, Mysterud Mysterud, Tverdal Tverdal, Kumpula Kumpula, Nieminen Nieminen (2007)
Male phenotypic quality influences offspring sex ratio in a polygynous ungulateProc. Roy. Soc. Lond. Ser. B, 274
P. Grant, B. Grant (1992)
DEMOGRAPHY AND THE GENETICALLY EFFECTIVE SIZES OF TWO POPULATIONS OF DARWIN'S FINCHES'Ecology, 73
Bjørneraas Bjørneraas, Solberg Solberg, Herfindal Herfindal, Sæther Sæther (2009)
Large‐scale spatiotemporal variation in calf sex ratio in moose ( Alces alces )an effect of density-dependent decrease in maternal condition?, 97
Frankham Frankham (1995)
Effective population size/adult population size ratios in wildlifea review, 66
Milner‐Gulland Milner‐Gulland, Bukreevea Bukreevea, Coulson Coulson, Lushchekina Lushchekina, Kholodova Kholodova, Bekenov Bekenov, Grachev Grachev (2003)
Reproductive collapse in saiga antelope haremsNature, 422
J. Ginsberg, E. Milner‐Gulland (1994)
Sex‐Biased Harvesting and Population Dynamics in Ungulates: Implications for Conservation and Sustainable UseConservation Biology, 8
Phillip Fenberg, K. Roy (2008)
Ecological and evolutionary consequences of size‐selective harvesting: how much do we know?Molecular Ecology, 17
(1997)
Wildl. Res. 1 (Suppl.): 525–537
K. Røed, Ø. Holand, Martin Smith, H. Gjøstein, J. Kumpula, M. Nieminen (2002)
Reproductive success in reindeer males in a herd with varying sex ratioMolecular Ecology, 11
S. Engen, T. Ringsby, B. Sæther, R. Lande, H. Jensen, Magnar Lillegård, H. Ellegren (2007)
EFFECTIVE SIZE OF FLUCTUATING POPULATIONS WITH TWO SEXES AND OVERLAPPING GENERATIONS, 61
(2009)
Sex ratio variation in harvested moose Alces alces calves: reflecting the population calf sex ratio or caused by selective hunting
G. Caughley (1977)
Analysis of vertebrate populations
Law Law (2007)
Fisheries‐induced evolutionpresent status and future directions, 335
Solberg Solberg, Ringsby Ringsby, Sæther Sæther, Heim Heim (2002)
Biased adult sex ratio can affect fecundity in primiparous mooseWildl. Biol., 8
B. Sæther, R. Andersen, O. Hjeljord, Morten Heim (1998)
Ecological Correlates of Regional Variation in Life History of the Moose Alces AlcesEcology, 79
R. Law (2007)
Fisheries-induced evolution: present status and future directionsMarine Ecology Progress Series, 335
Sæther Sæther, Solberg Solberg, Heim Heim, Stacy Stacy, Jakobsen Jakobsen, Olstad Olstad (2004a)
Offspring sex ratio in moose Alces alces in relation to paternal agean experiment, 10
E. Olsen, E. Olsen, M. Heino, G. Lilly, M. Morgan, J. Brattey, B. Ernande, U. Dieckmann (2004)
Maturation trends indicative of rapid evolution preceded the collapse of northern codNature, 428
B. Sæther, S. Engen, E. Solberg (2001)
Optimal harvest of age-structured populations of moose Alces alces in a fluctuating environmentWildlife Biology, 7
Climate influences on a population dynamics
K. Røed, Ø. Holand, A. Mysterud, A. Tverdal, J. Kumpula, M. Nieminen (2007)
Male phenotypic quality influences offspring sex ratio in a polygynous ungulateProceedings of the Royal Society B: Biological Sciences, 274
S. Engen, R. Lande, B. Saether (2005)
Effective Size of a Fluctuating Age-Structured PopulationGenetics, 170
E. Solberg, B. Sæther, O. Strand, A. Loison (1999)
Dynamics of a harvested moose population in a variable environmentJournal of Animal Ecology, 68
R. Waples (2002)
Effective size of fluctuating salmon populations.Genetics, 161 2
(2005)
Variation in male
R. Frankham (1995)
Effective population size/adult population size ratios in wildlife: a review.Genetical research, 89 5-6
J. Felsenstein (1971)
Inbreeding and variance effective numbers in populations with overlapping generations.Genetics, 68 4
W. Hill (1972)
Effective size of populations with overlapping generations.Theoretical population biology, 3 3
B. Sæther, S. Engen, E. Solberg, Morten Heim (2007)
Estimating the growth of a newly established moose population using reproductive valueEcography, 30
E. Milner‐Gulland, O. Bukreeva, Tim Coulson, A. Lushchekina, M. Kholodova, A. Bekenov, I. Grachev (2003)
Conservation: Reproductive collapse in saiga antelope haremsNature, 422
E. Solberg, M. Garel, Morten Heim, V. Grøtan, B. Sæther (2008)
Lack of compensatory body growth in a high performance moose Alces alces populationOecologia, 158
T. Coulson, F. Guinness, J. Pemberton, T. Clutton‐Brock (2004)
The demographic consequences of releasing a population of red deer from cullingEcology, 85
Sæther Sæther, Solberg Solberg, Heim Heim (2003)
Effects of altering sex ratio structure on the demography of an isolated moose populationJ. Wildl. Mgmt., 67
H. Jachmann, P. Berry, H. Imae (1995)
Tusklessness in African elephants: A future trendAfrican Journal of Ecology, 33
N. Ryman, R. Baccus, C. Reuterwall, Michael Smith (1981)
Effective Population Size, Generation Interval, and Potential Loss of Genetic Variability in Game Species under Different Hunting RegimesOikos, 36
B. Sæther (1997)
Environmental stochasticity and population dynamics of large herbivores: a search for mechanisms.Trends in ecology & evolution, 12 4
M. Festa‐Bianchet (2005)
Exploitative Wildlife Management as a Selective Pressure for Life-History Evolution of Large Mammals
A. Mysterud, T. Coulson, N. Stenseth (2002)
The role of males in the dynamics of ungulate populationsJournal of Animal Ecology, 71
Holand Holand, Røed Røed, Mysterud Mysterud, Kumpula Kumpula, Nieminen Nieminen, Smith Smith (2003)
The effect of sex ratio and male age structure on reindeer calvingJ. Wildl. Mgmt., 67
Charles Darwin (1930)
The Genetical Theory of Natural SelectionNature, 126
Kari Bjørneraas, E. Solberg, I. Herfindal, B. Sæther (2009)
Large-scale spatiotemporal variation in calf sex ratio in moose (Alces alces): an effect of density-dependent decrease in maternal condition?Canadian Journal of Zoology, 87
L. Nunney (1993)
THE INFLUENCE OF MATING SYSTEM AND OVERLAPPING GENERATIONS ON EFFECTIVE POPULATION SIZEEvolution, 47
(1969)
Evolution and the genetics of populations: the theory of gene
B. Sæther, E. Solberg, Morten Heim, J. Stacy, K. Jakobsen, Randi Olstad (2004)
Offspring sex ratio in moose Alces alces in relation to paternal age: an experimentWildlife Biology, 10
(1987)
Pattern and processes in the population dynamics of Scandinavian moose (Alces alces): some suggestions. Swed
Torunn Moe, E. Solberg, I. Herfindal, B. Sæther, Kari Bjørneraas, Morten Heim (2009)
Sex ratio variation in harvested moose (Alces alces) calves: does it reflect population calf sex ratio or selective hunting?European Journal of Wildlife Research, 55
Moe Moe, Solberg Solberg, Herfindal Herfindal, Sæther Sæther, Bjørneraas Bjørneraas, Heim Heim (2009)
Sex ratio variation in harvested moose Alces alces calvesreflecting the population calf sex ratio or caused by selective hunting?
R. Law (1979)
Harvest Optimization in Populations with Age DistributionsThe American Naturalist, 114
G. Proaktor, T. Coulson, E. Milner‐Gulland (2007)
Evolutionary responses to harvesting in ungulates.The Journal of animal ecology, 76 4
(2009)
488-495 c 2009 The Authors
Haldane Haldane (1927)
A mathematical theory of natural and artificial selection. V. Selection and mutationProc. Camb. Philos. Soc., 23
K. Røed, Ø. Holand, H. Gjøstein, H. Hansen (2005)
VARIATION IN MALE REPRODUCTIVE SUCCESS IN A WILD POPULATION OF REINDEER, 69
(1969)
Evolution and the genetics of populations: the theory of gene frequencies
F. Allendorf, P. England, G. Luikart, P. Ritchie, N. Ryman (2008)
Genetic effects of harvest on wild animal populations.Trends in ecology & evolution, 23 6
S. Lavsund, T. Nygrén, E. Solberg (2003)
Status of moose populations and challenges to moose management in Fennoscandia.Alces, 39
A. Lee, S. Engen, B. Saether (2008)
Understanding mating systems: a mathematical model of the pair formation process.Theoretical population biology, 73 1
E. Solberg, Morten Heim, V. Grøtan, B. Sæther, M. Garel (2007)
Annual variation in maternal age and calving date generate cohort effects in moose (Alces alces) body massOecologia, 154
S. Wright (1931)
Evolution in Mendelian Populations.Genetics, 16 2
(2004)
Offspring sex ratio in moose
B. Sæther, W. Sutherland, S. Engen (2004)
Climate Influences on avian population dynamicsAdvances in Ecological Research, 35
L. Slobodkin (1974)
Prudent Predation Does Not Require Group SelectionThe American Naturalist, 108
E. Pollak (1979)
Effective population numbers and mean times to extinction in dioecious populations with overlapping generations.Bellman Prize in Mathematical Biosciences, 52
D. Coltman, P. O'Donoghue, J. Jorgenson, J. Hogg, C. Strobeck, M. Festa‐Bianchet (2003)
Undesirable evolutionary consequences of trophy huntingNature, 426
R. Arthur (1960)
ON THE RELATION BETWEEN REPRODUCTIVE VALUE AND OPTIMAL PREDATION.Proceedings of the National Academy of Sciences of the United States of America, 46 1
(2003)
Reproductive collapse in saiga antelope
Sæther Sæther (1997)
Environmental stochasticity and population dynamics of large herbivoresa search for mechanisms, 12
A. Kuparinen, J. Merilä (2007)
Detecting and managing fisheries-induced evolution.Trends in ecology & evolution, 22 12
W. Ewens (1968)
Selection and Mutation
N. Leader‐Williams, R. Hudson, K. Drew, L. Baskin (1991)
Wildlife Production Systems: Economic Utilisation of Wild UngulatesJournal of Animal Ecology, 60
Holand Holand, Mysterud Mysterud, Røed Røed, Coulson Coulson, Gjøstein Gjøstein, Weladji Weladji, Nieminen Nieminen (2006)
Adaptive adjustment of offspring sex ratio and maternal reproductive effort in an iteroparous mammalProc. Roy. Soc. Lond. Ser. B, 273
L. Nunney, D. Elam (1994)
Estimating the effective population size of conserved populationsConservation Biology, 8
I. Herfindal, B. Saether, E. Solberg, R. Andersen, K. Høgda (2006)
Population characteristics predict responses in moose body mass to temporal variation in the environment.The Journal of animal ecology, 75 5
(2000)
Fishing as an evolutionary force
E. Solberg, V. Grøtan, C. Rolandsen, H. Brøseth, S. Brainerd (2005)
Change-in-sex ratio as an estimator of population size for Norwegian moose Alces alces, 11
Ø. Holand, A. Mysterud, K. Røed, T. Coulson, H. Gjøstein, R. Weladji, M. Nieminen (2006)
Adaptive adjustment of offspring sex ratio and maternal reproductive effort in an iteroparous mammalProceedings of the Royal Society B: Biological Sciences, 273
Sæther Sæther (1987)
Pattern and processes in the population dynamics of Scandinavian moose ( Alces alces )some suggestions, 1
Pollak Pollak (2000)
Effective population numbers and mean times to extinction in dioecious populations with overlapping generationsMath. Biosci., 52
The harvest of ungulate populations is often directed against certain sex or age classes to maximize the yield in terms of biomass, number of shot animals or number of trophies. Here we examine how such directional harvest affects the effective size of the population. We parameterize an age‐specific model assumed to describe the dynamics of Fennoscandian moose. Based on expressions for the demographic variance for a small subpopulation of heterozygotes Aa bearing a rare neutral allele a, we use this model to calculate how different harvest strategies influence the effective size of the population, given that the population remains stable after harvest. We show that the annual genetic drift, determined by , increases with decreasing harvest rate of calves and increasing sex bias in the harvest towards bulls 1 year or older. The effective population size per generation decreased with reduced harvest of calves and increased harvest of bulls 1 year or older. The magnitude of these effects depends on the age‐specific pattern of variation in reproductive success, which influences the demographic variance. This shows that the choice of harvest strategy strongly affects the genetic dynamics of harvested ungulate populations.
Animal Conservation – Wiley
Published: Oct 1, 2009
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